Habitus: Body yellow/orange (after preparation colourless or only slightly yellow). Eyes absent (also in living specimens). Cuticle: Dorso-lateral cuticle covered with a faint reticular pattern with irregular ridges, better developed caudally (mesh 1.7-5.1 µm in diameter) and with nine transverse rows of gibbosities (IX:4-6-5-6-5-6-4-2-2). Gibbosity rows II, IV and VI are in line with legs I, II and III, respectively. Lateral gibbosities in rows I and VII (i.e. gibbosities Ib and VIIb, respectively) shifted caudally relative to the dorsal gibbosities. In the remaining rows gibbosities arranged linearly. All legs without gibbosities but outer portions covered with sculpture. Ventral cuticle smooth (i.e., without sculpturing, tubercles or gibbosities). Buccal apparatus: Mouth antero-ventral. Peribuccal lamellae and papulae absent. Bucco-pharyngeal apparatus of the Macrobiotus type. Oral cavity armature comprises six ventral large round teeth arranged in two rows of three teeth each. However, in some specimens additional 1-5 smaller teeth are present in between the two main rows (e.g. in the holotype a there is single additional tooth in between the main median teeth). Buccal tube with a ventral lamina and with a single anterior bend (visible in lateral view only). In pharyngeal bulb two rod-shaped macroplacoids present, both without constrictions, length sequence 2<1. Microplacoid and septulum absent. Legs and claws: Claws of the Isohypsibius type, similar in shape on all legs. External claws I-III and anterior claws IV slightly larger than internal claws I-III and posterior claws IV, respectively. Primary branches of claws with thin accessory points ending at the highest point of the primary branch. Small, smooth and indistinct lunules present on all legs, but better developed only on legs IV. Bars and other cuticular thickenings on legs absent. Eggs: Four smooth eggs were found in one exuvium. Remarks: In some individuals both the cuticular sculpture and gibbosities are well defined whereas in others weakly developed and therefore difficult to identify under PCM.

Zawierucha, K., Michalczyk, Ł. & Kaczmarek, Ł. (2012) The first record of Tardigrada from the Republic of Zambia, with a description of Doryphoribius niedbalai sp. nov. (Eutardigrada: Isohypsibiidae, the evelinae group). African Zoology.