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  Animalia
      Tardigrada
            Eutardigrada
                    Parachela
                          Eohypsibioidea
                            Eohypsibiidae
                                     Amphibolus
                                     Austeruseus
                                             Austeruseus balduri
                                             Austeruseus faeroensis
                                             Austeruseus rokuri
                                     Bertolanius
                                             Bertolanius birnae
                                             Bertolanius mahunkai
                                                   [Amphibolus mahunkai]
                                                   [Macrobiotus mahunkai]
                                             Bertolanius markevichi
                                                   [Amphibolus markevichi]
                                             Bertolanius nebulosus
                                                   [Amphibolus nebulosus]
                                             Bertolanius portucalensis
                                             Bertolanius smreczynskii
                                                   [Amphibolus smreczynskii]
                                                   [Hypsibius smreczynskii]
                                                   [Isohypsibius smreczynskii]
                                             Bertolanius volubilis
                                                   [Amphibolus volubilis]
                                                   [Hypsibius volubilis]
                                                   [Isohypsibius volubilis]
                                             Bertolanius weglarskae
                                                   [Amphibolus weglarskae]
                                                   [Isohypsibius weglarskae]
                                     Eohypsibius
                                             Eohypsibius nadjae
                                             Eohypsibius terrestris